Patrice Francour - Full Professor
Fish, non indigenous species, marine protected areas
+33 (0)6 77 17 63 36
+33 (0)4 92 07 68 49

Specialist of fish populations in rocky areas and seagrass beds (Posidonia oceanica).My work takes place mainly in the marine protected areas (Port-Cros National Park, Marine Reserve of Scandola, Fishery reserve of Cantonment Cape Roux - Saint Raphael) and aims to understand the functioning of fish populations and the role of biodiversity in ecosystem functioning as well as the importance of habitat structure. I have developed new methods of in situ monitoring of fish populations, especially in the field of citizen science (FAST method). I regularly publish new or updated records of fish species for France or the Eastern Mediterranean (Gobiidae, Kyphosidae, Siganidae, Fistularidae, Oplegnathidae). One part of this activity is devoted to the non-indigenous species of fish spreading in the North-Western Mediterranean. In parallel, I have developed expertises to assist marine protected areas managers facing human activities (fishing, anchoring, etc). I recently carried out researchs involving Ecopath and Ecosim models (EwE) to analyze the role of the high trophic level predators in coastal ecosystems, particularly in relation to MPA management. All this work takes place in the western Mediterranean, but also in the eastern Mediterranean (Turkey in particular).



I. 99    Cuadros A., Moranta J., Cardona L., Thiriet P., Francour P., Vidal E., Sintes J., Cheminée A. 2019. Juvenile fish in Cystoseira forests: influence of habitat complexity and depth on fish behaviour and assemblage composition. Mediterranean Marine Science, 20(2). doi: 10.12681/mms.18857
I.98    Sreekanth G.B., Chakraborty S.K. Jaiswar A.K., Zacharia P.U. Mohamed K.S., Francour P. 2019. Trophic network and food web characteristics in a small tropical monsoonal estuary: A comparison with other estuarine systems. Indian Journal of Geo-Marine Sciences, in press.
I.97    Giakoumi S., Pey A., Di Franco A., Francour P., Kizilkaya Z., Arda Y., Raybaud V., Guidetti P. 2019. Exploring the relationships between marine protected areas and invasive fish in the world's most invaded sea. Ecological Applications, 29(1): e01809.



I.96    Ben Lamine E., Di Franco A., Romdhane M.S., Francour P. 2018. Comparing commercial, recreational and illegal coastal fishery catches and their economic values: A survey from the southern Mediterranean Sea. Fisheries Management and Ecology, 25: 456-463.
I.95    Riera E., Lamy D., Goulard C., Francour P., Hubas C. 2018. Biofilm monitoring as a tool to assess the efficiency of artificial reefs as substrates: toward 3D printed reefs. Ecological Engineering, 120: 230-237.
I.94    Taneez M., Hurel C., Mady F., Francour P. 2018. Capping of marine sediments with valuable industrial by-products: evaluation of inorganic pollutants immobilization. Environmental Pollution, 239: 714-721.
I.93     Vandenbussche P.S.P., Spennato G., Francour P., Pierson P.M. 2018. Validation of a simple and well-suited chemical cleaning method for fish otoliths. Acta Zoologica, 99(3): 258-262.
I.92     Rouane-Hacene O., Boutiba Z., Benaissa M., Belhaouari B., Francour P., Guibbolini-Sabatier M.E., Risso-De Faverney C. 2018. Seasonal assessment of biological indices, bioaccumulation, and bioavailability of heavy metals in sea urchins Paracentrotus lividus from Algerian west coast, applied to environmental monitoring. Environmental Science and Pollution Research, 25(12): 11238-11251.
I.91    Ben Lamine E., Romdhane M.S., Di Franco A., Francour P. 2018. Can citizen science contribute to fish assemblages monitoring in understudied areas? The case study of Tunisian marine protected areas. Estuarine, Coastal and Shelf Science, 200: 420-427.
I.90    Ben Lamine E., Guidetti P., Romdhane M.S., Francour P. 2018. Fish assemblages along the coasts of Tunisia: a baseline study to assess the effectiveness of future Marine Protected Areas. Mediterranean Marine Science, 19/1: 11-20.



I.89    Mouine-Oueslati N., Ines C., Ahlem R., Ktari M.H., Francour P. 2017. First biological data on the well established Lessepsian migrant Fistularia commersonii (Fistulariidae) in the Gulf of Tunis (Central Mediterranean). Russian Journal of Marine Biology, 43(6): 503-506.
I.88    Béarez P., Pruvost P., Feunteun E., Iglésias S., Francour P., Causse R., De Mazieres J., Tercerie S., Bailly N. 2017. Checklist of the marine fishes from metropolitan France. Cybium, 41(4): 351-371.
I.87    Cheminée A., Pastor J., Bianchimani O., Thiriet P., Sala E., Cottalorda J.M., Dominici J.M., Lejeune P., Francour P. 2017. Juvenile fish assemblages in temperate rocky reefs are shaped by the presence of macro-algae canopy and its three-dimensional structure. Scientific Reports, 7: 14638.
I.86     Prato G., Thiriet P., Di Franco A., Francour P. 2017. Enhancing fish underwater visual census to move forward assessment of fish assemblages: an application in three Mediterranean marine protected areas. PLoS ONE, 12(6): e0178511.
I.85    Tiquio M.G.J., Hurel C., Marmier N., Taneez M., Andral B., Jordan N., Francour P. 2017. Sediment-bound trace metals in Golfe-Juan Bay, northwestern Mediterranean: Distribution, availability and toxicity. Marine Pollution Bulletin, 118: 427-436..
I.84 Tiquio M.G.J., Marmier N., Francour P. 2017. Management frameworks for coastal and marine pollution in the European and South East Asian regions. Ocean and Coastal Management, 135: 65-78.



I.83 Piasecki W., G»abi½ski Z., Francour P., Koper P., Saba G., García A.M., Ünal V., Karachle P.K., Lepetit A., Tservenis R., K2z2lkaya Z., Stergiou K.I. 2016. Pescatourism - a European review and perspective. Acta Ichthyologica et Piscatoria, 46(4): 325-350.
I.82 Di Franco A., Thiriet P., Di Carlo G., Dimitriadis, C. Francour P., Gutiérrez N., Jeudy de Grissac A., Koutsoubas D., Milazzo M., Otero M., Piante C., Plass-Johnson J., Sainz-Trapaga S., Santarossa L., Tudela S., Guidetti P. 2016. Five key attributes can increase marine protected areas performance for small scale fisheries management. Scientific Report, 6:38135.
I.81 Prato G., Barrier C., Francour P., Cappanera V., Markantonatou V., Guidetti P., Mangialajo L., Cattaneo-Vietti R., Gascuel D. 2016. Assessing the interacting impacts of artisanal and recreational fisheries in a small Marine Protected Area (Portofino, NW Mediterranean Sea). Ecosphere7(12): e01601
I.80 Thiriet P.D., Di Franco A., Cheminée A., Guidetti P., Bianchimani O., Basthard-Bogain S., Cottalorda J.M., Arceo H., Moranta J., Lejeune P., Francour P., Mangialajo L. 2016. Abundance and diversity of crypto- and necto-benthic coastal fish are higher in marine forests than in structurally less complex macroalgal assemblages. PloS ONE, 11(10): e0164121.
I.79 Romdhani A., Ktari M.H., Dufour J.L., Mahe K., Francour P. 2016. Growth and age estimation of the greater forkbeard, Phycis blennoides (Actinopterygii: Gadiformes: Phycidae), from the Gulf of Tunis (central Mediterranean). Acta Ichthyologica et Piscatoria, 46 (1): 25-32.
I.78 Cheminée A., Merigot B., Vanderklift M.A., Francour P. 2016. Does habitat complexity influence fish recruitment? Mediterranean Marine Science, 17(1): 39-46.



I.77 Rouane-Hacene O., Boutiba Z., Belhaouari B., Guibbolini-Sabatier M.E., Francour P., Risso-de Faverney C. 2015. Seasonal assessment of biological indices, bioaccumulation and bioavailability of heavy metals in mussels Mytilus galloprovincialis from Algerian west coast, applied to environmental monitoring. Oceanologia, 57: 362-374.
I.76 Bussotti S., Di Franco A., Francour P., Guidetti P. 2015. Fish assemblages of Mediterranean marine caves. PlosOne, 10(4): e0122632.
I.75 Tessier A., Francour P., Charbonnel E., Dalias N., Bodilis P., Seaman W., Lenfant P. 2015. Assessment of French artificial reefs: due to limitations of research, trends may be misleading. Hydrobiologia, 753: 1-29.
I.74 Chater I., Romdhani A., Dufour J.L., Mahe K., Francour P., Chakroun-Marzouk N. 2015. Otolith growth and age estimation of bastard grunt, Pomadasys incisus (Actinopterygii: Perciformes: Haemulidae) in the Gulf of Tunis (Central Mediterranean). Acta Ichthyologica et Piscatoria, 45(1): 57-64.
I.73 Kara M.H., Ben Lamine E., Francour P. 2015. Range expansion of the invasive pufferfish, Lagocephalus sceleratus (Actinopterygii: Tetraodontiformes: Tetraodontidae) to the south-western Mediterranean. Acta Ichthyologica et Piscatoria, 45(1): 103-108.
I.72 Parravicini V., Mangialajo L., Mousseau L., Peirano A., Morri C., Montefalcone M., Francour P., Kulbicki M., Bianchi C.N. 2015. Climate change and warm-water species at border ecosystems in the Mediterranean Sea. Marine Ecology-An Evolutionary Perspective, doi: 10.1111


I.71 Prato G., Gascuel D. Valls A., Francour P. 2014. Balancing complexity and feasibility in Mediterranean coastal food-web models: uncertainty and constraints. Marine Ecology Progress Series, 512: 71-88.

I.70 Colléter M., Gascuel D., Albouy C., Francour P., Tito de Morais L., Valls A., Le Loc’h F. 2014. Fishing inside or outside? A case studies analysis of potential spillover effect from marine protected areas, using food web models. Journal of Marine Systems, 139: 383-395.

I.69 Marengo M., Durieux E.D.H., Marchand B., Francour P. 2014. A review of biology, fisheries and population structure of Dentex dentex. Reviews in Fish Biology and Fisheries, 24: 1065-1088.

I.68 Seytre C., Francour P. 2014. A long term survey of Posidonia oceanica fish assemblages in a Mediterranean marine protected area: emphasis on stability and no-take area effectiveness. Marine and Freshwater Research, 65(3): 244-254.
I.67 Bodilis P., Louisy P., Draman M., Francour P. 2014. Can citizen science survey non-indigenous fish species in the Eastern Mediterranean sea? Environmental Management, 53: 172-180.
I.66 Asnaghi V., Mangialajo L., Gattuso J.P., Francour P., Privitera D., Chiantore M. 2014. Effects of ocean acidification and diet on thickness and carbonate elemental composition of the test of juvenile sea urchins. Marine Environmental Research, 93: 78-84.


I.65 Gianni F., Bartolini F., Airoldi L., Ballesteros E., Francour P., Meinesz A., Thibaut T., Mangialajo L. 2013. Conservation and restoration of marine forests in the Mediterranean Sea and the potential role of Marine Protected Areas. Advances in Oceanography and Limnology, 4(2): 83-101.
I.64 Prato G., Guidetti P., Bartolini F., Mangialajo L., Francour P. 2013. The importance of high-level predators in Marine Protected Areas management: consequences of their decline and their potential recovery in the Mediterranean context. Advances in Oceanography and Limnology, 4(2): 176-193.
I.63 Tessier A., Pastor J., Francour P., Saragoni G., Crec'hriou R., Lenfant P. 2013. Video transect as a complement to underwater visual census to study reserve effect on fish assemblages. Aquatic Biology, 18: 229-241.
I.62 Asnaghi V., Chiantore M., Mangialajo L., Gazeau F., Francour P., Alliouane S., Gattuso J.P. 2013. Cascading Effects of Ocean Acidification in a Rocky Subtidal Community. PlosOne, 8(4): e61978.
I.61 Seytre C., Vanderklift M., Bodilis P., Cottalorda J.M., Gratiot J., Francour P. 2013. Assessment of commercial and recreational fishing effects on trophic interactions in the Cap Roux area (north western Mediterranean). Aquatic Conservation: Marine and Freshwater Ecosystems, 23: 189-201.
I.60 Bodilis P., Crocetta F., Langeneck J., Francour P. 2013. The spread of an Atlantic fish species, Pomadasys incisus (Bowdich, 1825) (Osteichthyes: Haemulidae), within the Mediterranean Sea with new additional records from the French Mediterranean coast. Italian Journal of Zoology, 80(2): 273-278.
I.59 Cheminée A., Sala E., Pastor J., Bodilis P., Thiriet T., Mangialajo L., Cottalorda J.M., Francour P. 2013. Nursery value of Cystoseira forests for Mediterranean rocky reef fishes. Journal of experimental Marine Biology and Ecology, 442: 70-79.
I.58 Arceo H., Cazalet B., Aliño P., Mangialajo L., Francour P. 2013. Moving beyond a top-down fisheries management approach in the northwestern Mediterranean: some lessons from the Philippines. Marine Policy, 39: 29-42.


I.57 Mezali K., Francour P. 2012. Les holothuries Aspidochirotes de quelques sites des côtes algériennes : révision systématique et relations phylogénétiques. Bull. Soc. zool. France, 137(1-4): 177-192.
I.56 Chakroun N., Houman B., Benaissa Z., Francour P. 2012. Situation alarmante des aires protégées tunisiennes pendant la révolution : Échec d’une politique de conservation et remèdes. Quid de la recherche ?. Bull. Soc. zool. France, 137(1-4): 29-43.
I.55 Bodilis P., Arceo H., Cheminée A., Miniconi R., Francour P. 2012. Occurrences of the Atlantic species, Pisodonophis semicinctus (Osteichthyes: Ophichthidae), along the Mediterranean coast. Journal of Applied Ichthyology, 28(4):652-654.
I.54 Valls A., Gascuel D., Guénette S., Francour P. 2012. Modeling trophic interactions to assess the potential effects of a marine protected area: case study in the NW Mediterranean Sea. Marine Ecology Progress Series, 456: 201-214.
I.53 Sartoretto S., Francour P. 2012. Bathymetric distribution and growth rates of Eunicella verrucosa (Cnidaria: Gorgoniidae) populations along the Marseilles coast (France). Scientia marina, 76(2): 349-355.
I.52 Mouine N., Francour P., Ktari M.H., Chakroun-Marzouk N. 2012. Reproductive biology of four Diplodus species Diplodus vulgaris, D. annularis, D. sargus sargus and D. puntazzo (Sparidae) in the Gulf of Tunis (central Mediterranean). Journal of the Marine Biological Association UK, 92(3): 623-631.


I.51 Forcioli D., Merle P.L., Caligara C., Ciosi M., Muti C., Francour P., Cerrano C., Allemand D. 2011. Symbiont diversity is not involved in depth acclimation in the Mediterranean sea whip Eunicella singularis. Marine Ecology Progress Series, 439: 57-71.
I.50 MerMex Group, Francour P. (parmi 95 auteurs classés par ordre alphabétique) 2011. Marine ecosystems’ responses to climatic and anthropogenic forcings in the Mediterranean. Progress in Oceanography, 91: 97-166.
I.49 Bodilis P., Seytre C., Charbonnel E., Francour P. 2011. Monitoring of the fish assemblages of the artificial reefs of Golfe Juan marine protected area (France, the north-western Mediterranean). Brazilian Journal of Oceanography, 59: 167-176.
I.48 Cheminée A., Francour P., Harmelin-Vivien M. 2011. Assessment of Diplodus spp. (Sparidae) nursery grounds along Marseilles rocky shore (France, NW Mediterranean). Scientia marina, 75(1): 181-188.


I.47 Pastor J., Francour P. 2010. Occurrence and distribution range of Parablennius pilicornis (Cuvier, 1829) (Actinopterygii: Perciformes: Blenniidae) along the French Mediterranean coast. Acta Ichthyologica et Piscatoria, 40(2): 179-185.
I.46 Schrembi P.J., Bodilis P., Evans J., Francour P. 2010. Occurrence of barred knifejaw, Oplegnathus fasciatus (Actinopterygii: Perciformes: Oplegnathidae), in Malta (Central Mediterranean) with a discussion on possible modes of entry. Acta Ichthyologica et Piscatoria, 40(2): 101-104.
I.45 Risso-de Faverney C., Guibbolini-Sabatier M.E., Francour P. 2010. An ecotoxicological approach with transplanted mussels (Mytilus galloprovincialis), for assessing the impact of tyre reefs immersed along the NW Mediterranean Sea. Marine Environmental Research, 70: 87-94.
I.44 Francour P., Cottalorda J.M., Aubert M., Bava S., Colombey M., Gilles P., Kara H., Lelong P., Mangialajo L., Miniconi R., Quignard J.P. 2010. Recent occurrences of Opah Lampris guttatus (Lampridae) in the western Mediterranean Sea. Acta Ichthyologica et Piscatoria, 40(1): 91-98.
I.43 Lassauque J., Lepoint G., Thibaut T., Francour P., Meinesz A. 2010. Tracing sewage and natural freshwater input in a Northwest Mediterranean bay: evidence obtained from isotopic ratios in marine organisms. Marine Pollution Bulletin, 60: 843-851.


I.42 Daniel B., Piro S., Charbonnel E., Bachet F., Francour P., Letourneur Y. 2009. Lessepsian rabbitfish Siganus luridus reached the French Mediterranean coasts. Cybium, 33(2): 163-164.
I.41 Francour P., Pellissier V., Mangialajo L., Buisson E., Stadelmann B., Veillard N., Meinesz A., Thibaut T., Vaugelas de V. 2009. Changes in invertebrate assemblages of Posidonia oceanica beds following Caulerpa taxifolia invasion. Vie Milieu, 59(1): 31-38.
I.40 Seytre C., Francour P. 2009. The Cap Roux MPA (Saint-Raphaël, French Mediterranean): changes in fish assemblages within four years of protection. ICES Journal of Marine Science, 66: 180-187.


I.39 Francour P. 2008. First records of Didogobius splechtnai along the French Mediterranean coast and additional comments about D. schlieweni. Acta Ichthyologica et Piscatoria, 38(2): 139-142.
I.38 Seytre C., Francour P. 2008. Is the Cape Roux marine protected area (Saint-Raphaël, Mediterranean Sea) an efficient tool to sustain artisanal fisheries ? First indications from visual censuses and trammel net sampling. Aquatic Living Resources, 21(3): 297-305.
I.37 Francour P., Mouine N. 2008. First record of Kyphosus sectator (Linnaeus, 1758) (Kyphosidae) along the French Mediterranean coast. Cybium, 32(3): 275-276.


I.36 Francour P., Mangialajo L. 2007. Gobius kolombatovici, a common species of Gobiidae in the north-western Mediterranean Sea ? Cybium, 31(3): 389-390.
I.35 Mouine N., Francour P., Ktari M.H., Chakroun-Marzouk N. 2007. The reproductive biology of Diplodus sargus sargus in the Gulf of Tunis (central Mediterranean). Scientia Marina, 71(3): 461-469.
I.34 Dufour F., Guidetti P., Francour P. 2007. Comparaison des inventaires de poissons dans les aires marines protégées de Méditerranée : influence de la surface et l'ancienneté. Cybium, 31(1): 19-31.


I.33 Casagranda C., Boudouresque C.F., Francour P. 2005. Abundance, population structure and production of Hydrobia ventrosa (Gastropoda: Prosobranchia) in a Mediterranean brackish lagoon, Lake Ichkeul, Tunisia. Archiv für Hydrobiologie, 164(3): 411-428.
I.32 Pergent-Martini C., Leoni V., Pasqualini V., Ardizzone G. D., Balestri E., Bedini R., Belluscio A., Belsher T., Borg J., Boudouresque C. F., Boumaza S., Bouquegneau J. M., Buia M. C., Calvo S., Cebrian J., Charbonnel E., Cinelli F., Cossu A., Di Maida G., Dural B., Francour P., Gobert S., Lepoint G., Meinesz A., Molenaar H., Mansour H. M., Panayoditis P., Peirano A., Pergent G., Piazzi L., Pirrotta M., Relini G., Romero J., Sanchez-Lizaso J. L., Semroud R., Shembri P., Shili A., Tomasello A., Velmirov B. 2005. Descriptors of Posidonia oceanica meadows: use and application. Ecological Indicators, 5: 213-230.
I.31 Longepierre S., Robert A., Levi F., Francour P. 2005. How invasive alga species (Caulerpa taxifolia) induces changes in foraging strategies of the benthivorous fish Mullus surmuletus in coastal Mediterranean ecosystems. Biodiversity and Conservation, 14: 365-376.


I. 30 Levi F., Francour P. 2004. Behavioural response of Mullus surmuletus to habitat modification by the invasive macroalga Caulerpa taxifolia. Journal of Fish Biology, 64: 55-64.
I.29 Francour P., Bodilis P. 2004. La protection des peuplements de poissons de la réserve naturelle de Scandola (Corse) influence-t-elle les performances de l’écosystème ? Revue d’Ecologie (Terre et Vie), 59: 363-365.


I.28 Bodilis P., Ganteaume A., Francour P. 2003. Recruitment of the dusky grouper (Epinephelus marginatus) in the north-western Mediterranean sea. Cybium, 27(2): 123-129.
I.27 Francour P., Javel F. 2003. Recent occurrences of young Schedophilus ovalis (Centrolophidae) along French Mediterranean coasts. Cybium, 27(1): 57-58.
I.26 Bodilis P., Ganteaume A., Francour P. 2003. Presence of 1 year-old dusky groupers along the French Mediterranean coast. Journal of Fish Biology, 62: 242-246.


I.25 Arigoni S., Francour P., Harmelin-Vivien M., Zaninetti L. 2002. Adaptive coloration of Mediterranean labrid fishes to the new habitat provided by the introduced tropical alga Caulerpa taxifolia. Journal of Fish Biology, 60: 1486-1497.


I.24 Francour P., Harmelin J.G., Pollard D., Sartoretto S. 2001. A review of marine protected areas in the north-western Mediterranean region: sitting, usage, zonation and management. Aquatic Conservation: Marine and Freshwater Ecosystems, 11: 155-188.
I.23 Soltan D., Verlaque M., Boudouresque C.F., Francour P. 2001. Changes in macroalgal communities in the vicinity of a Mediterranean sewage outfall after the setting up of a treatment plant. Marine Pollution Bulletin, 42(1): 59-70.


I.22 Francour P. 2000. Evolution spatio-temporelle à long terme des peuplements de poissons des herbiers à Posidonia oceanica de la réserve naturelle de Scandola (Corse, Méditerranée nord-occidentale). Cybium, 24(3) suppl.: 85-95.
I.21 Pinnegar J.K., Polunin N.V.C., Francour P., Badalamenti F., Chemello R., Harmelin-Vivien M., Hereu B., Milazzo M., Zabala M., D’Anna G., Pipitone C. 2000. Trophic cascades in benthic marine ecosystems: lessons for fisheries and protected-area management. Environmental Conservation, 27(2): 179-200.
I.20 Pérez T., Garrabou J., Sartoretto S., Harmelin J.G., Francour P., Vacelet J. 2000. Mortalité massive d’invertébrés marins : un événement sans précédent en Méditerranée nord-occidentale. Comptes Rendus Académie des Sciences, Sciences de la Vie, 323: 853-865.
I.19 Guettaf M., San Martin G.A., Francour P. 2000. Interpopulation variability of the reproductive cycle of Paracentrotus lividus (Echinodermata: Echinoidea) in the south-western Mediterranean. Journal of the Marine Biological Association UK, 80: 899-907.
I.18 Ruitton S., Francour P., Boudouresque C.F. 2000. Relationships between algae, benthic herbivorous invertebrates and fishes in rocky sublittoral communities of a temperate sea (Mediterranean). Estuarine, Coastal and Shelf Science, 50: 217-230.


I.17 Francour P., Ganteaume A., Poulain M. 1999. Effect of boat anchoring in Posidonia oceanica seagrass beds in the Port-Cros national park (north-western Mediterranean sea). Aquatic Conservation: Marine and Freshwater Ecosystems, 9: 391-400.


I.16 Ganteaume A., Gobert J., Malestroit P., Ménager V., Francour P., Boudouresque C.F. 1998. In vitro consumption of Caulerpa taxifolia by Paracentrotus lividus: seasonal variation and tolerance effect. Journal of the Marine Biological Association UK, 78: 239-248.


I.15 Sartoretto S., Francour P., Harmelin J.G., Charbonnel E. 1997. Observations in situ de deux Labridae profonds, Lappanella fasciata (Cocco, 1833) et Acantholabrus palloni (Risso, 1810), en Méditerranée nord-occidentale. Cybium, 21(1): 37-44.
I.14 Sartoretto S., Francour P. 1997. Quantification of bioerosion by Sphaerechinus granularis on "coralligene" concretions of the western Mediterranean. Journal of the Marine Biological Association UK, 77: 565-568.
I.13 Macpherson E., Biagi F., Francour P., Garcia-Rubies A., Harmelin J.G., Harmelin-Vivien M., Jouvenel J.Y., Planes S., Vigliola L., Tunesi L. 1997. Mortality of juvenile fishes of the genus Diplodus in protected and unprotected areas in the western Mediterranean sea. Marine Ecology Progress Series, 160: 135-147.
I.12 Francour P. 1997. Fish assemblages of Posidonia oceanica beds at Port-Cros (France, NW Mediterranean): assessment of composition and long term fluctuations by visual census. P.S.Z.N.I.: Marine Ecology, 18(2): 157-173.
I.11 Francour P. 1997. Predation on holothurians: a literature review. Invertebrate Biology, 116(1): 52-60.


I.10 Francour P., Thibault J.C. 1996. The diet of breeding osprey Pandion haliaetus on Corsica: exploitation of a coastal marine environment. Bird Study, 43: 129-133.


I.9 Francour P., Harmelin-Vivien M., Harmelin J.G., Duclerc J. 1995. Impact of Caulerpa taxifolia colonization on the littoral ichthyofauna of North-Western Mediterranean sea: preliminary results. Hydrobiologia, 300/301: 345-353.


I.8 Francour P., Boudouresque C.F., Harmelin J.G., Harmelin-Vivien M., Quignard J.P. 1994. Are the Mediterranean waters becoming warmer ? Information from biological indicators. Marine Pollution Bulletin, 28(9): 523-526.
I.7 Francour P. 1994. Pluriannual analysis of the reserve effect on ichtyofauna in the Scandola natural reserve (Corsica, northwestern Mediterranean). Oceanologica Acta, 17(3): 309-317.


I.6 Harmelin-Vivien M.L., Francour P. 1992. Trawling or visual censuses ? Methodological bias in the assessment of fish populations in seagrass beds. P.S.Z.N.I. Marine Ecology, 13(1): 41-51.
I.5 Francour P., Semroud R. 1992. Calculation of the root area index in Posidonia oceanica in the western Mediterranean. Aquatic Botany, 42: 281-286.


I.4 Francour P. 1991. The effect of protection level on a coastal fish community at Scandola, Corsica. Revue d’Ecologie (Terre et Vie), 46 : 65-81.


I.3 Miniconi R., Francour P., Bianconi C.H. 1990. Inventaire de la faune ichtyologique de la réserve naturelle de Scandola (Corse, Méditerranée nord-occidentale). Cybium, 14(1): 77-89.
I.2 Francour P., Marchessaux D., Argiolas A., Campredon P., Vuignier G. 1990. La population de phoque moine (Monachus monachus) de Mauritanie. Revue d’Ecologie (Terre et Vie), 45: 55-64.
I.1 Chauvet C., Francour P. 1990. Les mérous du parc national de Port-Cros : aspects socio-démographiques. Bull. Soc. zool. France, 114(4): 5-13.


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D.U. - Médecine de la plongée et des activités sub-aquatiques
    Cours : Les Espèces de faune et flore potentiellement dangereuses pour le plongeur

Conférences diverses

Cours LSVIII – Ecosystèmes 1° Semestre



Cours LSVIII – Biodiversité 2° Semestre

Ressources optionnelles Biodiversité


Cours LSVIII – Ecologie Marine 2° Semestre


Dynamiques des Populations 1° Semestre
    Cours 1: Méta-populations
        TD : Méta-populations sur Scandola       Simulations sous Excel
        TD : Fragmentation

    Cours 2 : Distribution des Individus en Espèces; Indices de Diversité
        TD : Indices de Diversité

    Cours 3 : Modélisation des réseaux trophiques
        TD : Modèles Ecopath

Publications sur ECOPATH
        Valls et al. 2012     Valls et al. 2012 suppléments
        Colléter et al. 2014
        Prato et al. 2014

    Rappels Dynamique des Populations
    Rappels Biodiversité et Ecosystèmes
    Classification des Espèces